Brownie is a program for analyzing rates of continuous character evolution on a tree or set of trees. See http://www.brianomeara.info/brownie for more info.

This version incorporates trait evolution and species delimitation methods.

Funding for its development has come from a University of California, Davis Population Biology fellowship, NSF Graduate Research Fellowship and Dissertation Improvement Grant (DEB-0608487) awards to Brian O'Meara, and from the National Evolutionary Synthesis Center (NESCent) (NSF #EF-0423641).